A distant Federation outpost Fort Casey comes under attack by bugs. The team on the fast attack ship Alesia is assigned to help the Starship John A. Warden stationed in Fort Casey evacuate along with the survivors and bring military intelligence safely back to Earth. Carl Jenkins, now ministry of Paranormal Warfare, takes the starship on a clandestine mission before its rendezvous with the Alesia and goes missing in the nebula. Now, the battle-hardened troopers are charged with a rescue mission that may lead to a much more sinister consequence than they ever could have imagined….

In the final months of World War II, 14-year-old Seita and his sister Setsuko are orphaned when their mother is killed during an air raid in Kobe, Japan. After a falling out with their aunt, they move into an abandoned bomb shelter. With no surviving relatives and their emergency rations depleted, Seita and Setsuko struggle to survive.

Federation trooper Johnny Rico is ordered to work with a group of new recruits on a satellite station on Mars, where giant bugs have decided to target their next attack.